Kyle Vantrease threw four touchdown passes, including a 22-yarder to Derwin Burgess Jr. with 1:10 remaining, and Georgia Southern beat No. 25 James Madison 45-38 to spoil the Dukes’ first appearance in the AP Top 25.

Dukes score 29 unanswered points for the road win over the Mountaineers.

Spencer Petras threw for 224 yards and two touchdowns and No. 5 Iowa needed a second-half comeback to defeat Colorado State 24-14.